Psammodes; Print; The Tok-tok beetles (genus Psammodes) are ground-dwelling; Afrotropical beetles in the family Tenebrionidae. They are stout in shape and typically black or dark rufous in colour. They average about 2.6 cm in body length; but the Spindle toktokkie (P. bertolonii) has an elongate body shape and measures 5.3 cm; while P. sulcicollis reaches 6 to 8 cm; and is the largest Tenebrionid in the world. Like the related genus Dichtha; the adults tap out a rhythm on the ground to attract and locate mates. Habitats are varied; from coastal forest to ridges; koppies; woodland and desert sand.